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Antagonistic microorganisms
Bacteria, fungi or other microorganisms that compete for food, by producing compounds inhibitory to other microorganisms or by directly parasitizing them.

Apex
The root tip covered by a root cap and secreted mucilage, i.e. soluble complex sugars (poly saccharides), which adhere to roots.

Apical root meristem
The zone of dividing cells at the apex which gives rise to new cells and eventually new tissues in a growing root cell.

Bacteria
Primitive, unicellular, microscopic organisms that lack a cell nucleus and other organelles, obtain soluble food by absorption, and reproduce by simple cell division.

Beneficial fungus
A fungus which has a beneficial effect on an important process essential to plant growth, ie. mycorrhizal fungi, litter decomposers, fungi antagonistic to pathogens, etc.

Biological control
A term used to describe the use of natural organisms to reduce the effects of undesirable organisms (pests) and to favor desirable organisms such as crops, trees, animals, beneficial organisms or insects.

Casparian strip
A subarin layer in endodermis that function as a selective barrier allowing absorption of desirable elements (N, P, K, Ca, etc.) but blocking absorption of harmful elements (Na, Al, etc.)

Cellulose
A polysaccharide (i.e., Many sugars) made up of many glucose molecules chemically bonded together. The most abundant organic compound on earth. It comprises the bulk of cell walls of plants where it occurs as microfibrils (i.e., fiber strands).

Chlorophyll
The green pigment of plant cells necessary for photosynthesis by capturing light energy and coverting it to chemical energy.

Colonization
The establishment of a species in a given environment.

Competition
The struggle among organisms for a limited supply of water, food, light, etc. in a given environment.

Competitors
Refers to two species of organism which compete with each other for survival by attempting to use the same substrate or by direct competition with each other.

Cortex
The cell layers occurring between the epidermis and the stele or vascular cylinder.

Ecosystem
A term used to describe a system of interactions between living organisms occupying the same environment.

Ectomycorrhizal fungi
Species of mycorrhizal fungi with hyphae that grow only between the cells and over the surface (outside) of the roots, but do not penetrate the plant cell.

Endodermis
The inner most layer of cortical cells ("inner skin") found in all roots next to the vascular cylinder. The endodermis contains a Casparian strip.

Endomycorrhizal fungi
Species of mycorrhizal fungi with hyphae that penetrate and live inside the cortical cells of the plant root.

Epidermis
The outer most layer of cells ("outer skin") of a growing root in direct contact with the soil.

Ericoid mycorrhizal fungi
Species of mycorrhizal fungi which exhibit some of the characteristics of both endo- and ectomycorrhizae, and are unique to azaleas, rhododendrons, blueberry, cranberry and eastern mountain laurel.

Fine absorbing roots
Tiny roots usually located in the upper several inches of soil that are actively engaged in the uptake of water and minerals.

Fungus
A non-photosynthetic, chiefly multicellular organism only able to use carbon, nitrogen, etc. as starting materials for synthesis of its essential foods.

Fungus mantle
A sheath made up of the hyphae of an ectomycorrhizal fungus which surrounds and encompasses the fine absorbing roots.

Host
An organism on or in a parasite lives.

Hypha (hyphae, pl.)
A single tubular filament of a fungus.

Hyphal strand
An aggregation of several individual hyphae into a thread-like feeding structure.

Indigenous
Existing or growing naturally in a region or country.

Inoculate
To bring one organism into contact with a host plant or plant organ.

Inoculum
That part of the organism which is brought into contact with the host or the host environment in order to establish it there. In the case of the mycorrhizal fungi, this consists of spores or hyphae of the beneficial fungus.

Lignin
Phenolic compounds impregnating and strengthening cell walls. Xylem cells and fibers are typically lignified but other cells in the stele or cortex can also be lignified.

Lipids:
Diverse class compounds, including fats, oils, fatty acids, triglycerides and steroids essential for membrane formation, energy stores and fuel molecules.

Macronutrients
Mineral elements required by plants in large amounts - nitrogen, phosphorus and potassium (N-P-K)

Micronutrients
Elements which are required in very small or trace amounts - iron, chlorine, copper, manganese, zinc, molybdenum, boron etc.

Mycelium
The mass of hyphae forming the body of a fungus.

Mycorrhiza (mycorrhizae, pl.)
"Fungus roots" or hyphae of certain fungi (mycorrhizal fungi) which are symbiotic with the plant roots on which they occur. The symbiotic combination of the hyphae and the plant roots.

Nitrogen fixation
The incorporation of atmospheric nitrogen into nitrogen compounds that are available for uptake by green plants, a process that can be carried out only by certain microorganisms or by higher plants in symbiotic association with microorganisms.

Nutrients
Refers to the mineral elements usually found in the soil which are essential for plant growth.

Organs
Tissues that act together to perform functions. The roots, stems and leaves are the major organs of plants. All organs are composed of three tissues…epidermis, vascular tissues and ground tissues. Ground tissues are those involved in metabolism (such as photosynthesis), food storage and physical support.

Parasite
An organism that lives on or in an organism of a different species and derives nutrients from it.

Pathogen
An organism that causes a disease.

Pericycle
Special cells in endodermis that produce lateral roots.

Perderm:
The bark layer formed on the surface of roots (or branches) by secondary growth. It is strenthened by subarin and lignin.

Phenolics
Acid ring (aromatic) structure, organic compound derived from other hydrocarbons (i.e. contain C, Hand O).

Photosynthesis
The process by which carbon dioxide and water are combined in the presence of light energy and chlorophyll to form carbohydrates.

Plant Health Care (PHC) Management
An integrated management approach to plant health coordinated by the International Society of Arboriculture, National Arborist Association and USDA Forest Service; PHC uses multiple tactics and relies primarily on preventive measures, those that encourage plant vigor and maintain healthy, balanced growth, rather than emphasizing chemical treatments for diseases and insects.

Pt
Letters used to designate Pisolithus tinctorius, a major ectomycorrhizal fungus.

Radical (primary root)
The initial root formed after seed germination. This primary (or first) root produces first-order lateral roots. The primary root eventually becomes the tap root of the plant.

Rhizosphere
The soil near a living root, usually applied to the zone around tiny fine absorbing roots.

Rhizosphere competent
Term used to designate the ability of a mycorrhizal fungus or rhizobacteria to successfully occupy the rhizosphere by growing and spreading onto new roots of the host plant as roots develop.

Root hairs
Thin hair-like extensions of epidermal cells formed behind the root meristem. They increase root contact with the soil and are important for water and mineral element uptake but are very short lived.

Root systems
Composed of primary and secondary roots and different orders of lateral roots. Different orders of lateral roots vary in there growth rates, life spans, structural features, etc. as well as there capacity to obtain water and mineral elements or to support mycorrhizae. Higher order lateral roots are generally thinner, shorter, have shorter life spans and grow less rapidly than those of lower orders.

Root tissues
These are produced by cell division in the root tip (apex) and cell expansion behind the apex (subapex) regions. The tissues are mainly cellulose and lignin built with photosynthates. These tissues progressively mature at greater distances from the root tip.

Saprophyte
An organism that uses dead organic material for food.

Spores
The reproductive unit of fungi, bacteria and other microbes consisting of one or more cells; it is analogous to the seed of green plants.

Sporulation
The production of spores.

Stele or vascular cylinder
This zone inside the endodermis contains specialized tissues responsible for transport of water and mineral elements to the shoot (xylem) and organic nutrients, such as photosynthetically-fixed carbon, to the roots (phloem). Xylem cells develop lignified walls and are dead when mature.

Suberin:
A hydrophobic (lacks affinity for water) material, containing lipids and phenolics, which impregnates the walls of specialized cells and prevents the passage of water and other materials in the apoplast.

Substrate
The material on which a microorganism feeds and develops.

Symbiosis
A mutually beneficial association of two different kinds of organisms.

Virulence
The degree of pathogenicity of a given pathogen.

VAM
Abbreviation for vesicular-arbuscular mycorrhizal fungi, the most common type of endomycorrhizae.

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